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WILD LOBAU
SPEEDWELL FAMILY

THE SPEEDWELL FAMILY (Plantaginaceae or Veronicaceae)

This is another flower family very difficult to describe in a satisfying way. The oldest name given in 1782 was Veronicaceae which would tally with the type flower taken here (Speedwell’s latin name being Veronica). The name Antirrhinaceae was also proposed but, in the meantime the Angiosperm Phylogeny Group has accepted the name Plantaginaceae. This choice seems puzzling to the lay botanist as the name is reminiscent of the member genus Plantago, the plantains which, with their small inconspicuous flowers, don’t appear typical. (These plantains have no connection with the banana plant, a different family altogether). To make things more confusing, many members of this family were formerly considered part of the Scrophulariaceae, the figwort family. The reasons for all these changes lie in the field of microbiological and molecular studies. The family is so cosmopolitan and diverse that it’s very difficult to describe it at all. They occur mostly in temperate zones and include herbs, shrubs and also a few aquatic plants with roots.

The web site of the Natural History Museum in Vienna, Botanik im Bild, uses the family description Veronicaceae so I have also included it in the title.

Leaves are very diverse including spiral to opposite arrangements and simple to compound shapes. The structure and form of the flowers is also variable. Some genera are 4-merous (i.e. with 4 sepals and 4 petals), others are 5-8-merous. The flowers are often two-lipped but the type flower, speedwell, is not. Fruits are usually capsules with seeds that are shaken out when dry.

(source: Wikipedia, Botanik im Bild, web site of the Natural History Museum, Vienna)

Click on a thumbnail photo to go to full photo and description.

Common Toadflax in Lobau Austrian Speedwell in Lobau Saw-leaved Speedwell in Lobau Persian Speedwell in Lobau
Lobau Location
QUICK KEY

Leaf Types (Simplified)

Icon
Shape
Veins
Simple parallel leaf-vein diagram
Simple
Parallel
Simple pinnate leaf-vein diagram
Simple
Pinnate
Lobed pinnate leaf-vein diagram
Lobed
Pinnate
Compound pinnate leaf-vein diagram
Compound
Pinnate
Trifoliate3-pinnate leaf-vein diagram
Trifoliate
3-Pinnate
Simple palmate leaf-vein diagram
Simple
Palmate
Lobed palmate leaf-vein diagram
Lobed
Palmate
Compound palmate leaf-vein diagram
Compound
Palmate
Other and various leaf-vein arrangments diagram
Other
Various

Flower Types (Simplified)

Icon
Symmetry
Petals
Radial symmetry and 3 petals flower type diagram
Radial
3 or multiple 3s
Radial symmetry and 4 petals flower type diagram
4
Radial symmetry and 5 separate petals flower type diagram
Radial
5, Separate
Radial symmetry and 5 tubular petals flower type diagram
Radial
5, Tubular
Radial symmetry and 5 tubular base flower type diagram
Radial
5,Tubular base
Radial symmetry and 5 bell-shape petals flower type diagram
Radial
5, Bell shape
Bilateral symmetry and 5 separate petals flower type diagram
Bilateral
5, Separate
Bilateral symmetry and 5 lipped petals flower type diagram
Bilateral
5, Lipped
Bilateral symmetry and winged petals flower type diagram
Bilateral
Butterfly form
Bilateral symmetry and tubular flower type diagram
Bilateral
Tubular
Spurred flower type diagram
.
Spurred
Stellate composite flower type diagram
Stellate
Composite
Structured clusters flower type diagram
Structured Clusters
5, small
Anomalous flower type diagram
Anomalous
.
Inconspicuous flower type diagram
Inconspicuous
.

Inflorescence Types (Simplified)

Icon
Type
Capitulum
Umbel flower head inflorescence type diagram
Umbel
Spike-like flower head inflorescence type diagram
Spike-like
Branched flower head inflorescence type diagram
Branched
Whorled flower head inflorescence type diagram
Whorled
Clustered flower head inflorescence type diagram
Clustered
Single
Common Toadflax
 in Lobau

© 2012 Margaret Eros

LEAF TYPE
PETAL TYPE
INFLORES- CENCE TYPE
HEIGHT OF PLANT
FLOWERING MONTH
HABITAT
PHOTO LOCATION
  • diagram of 
<!--Next: FOR BLIND LEAF SHAPE ICON.-->
simple 
<!--Next brown section: FOR BLIND LEAF VEIN ICON.-->
 leaf shape and 
pinnate 
vein structure
    • diagram of 
<!--Next: FOR BLIND POPUP LEAF SHAPE.-->
simple 
leaf shape 
<!--Next: FOR BLIND POPUP LEAF VEIN ICON.-->
and 
pinnate 
vein structure
    • LEAF TYPE
    • Shape:
    • --simple (undivided/entire).
    • Veins:
    • --pinnate (main central vein with side branches).
    • This plant:
    • Narrow, up to 60mm long, fine, soft and dense giving feathery appearance
  • Bilateral symmetry and lipped petals
flower type diagram
    • 
Bilateral symmetry and lipped petals
flower type diagram
    • FLOWER TYPE
    • Symmetry:
    • --bilateral (distinct right and left sided [mirror image])
    • Petals:
    • --lipped (lower petal forms distinct lip with platform, upper ones may form a ‘hood’).
    • This plant:
    • Long straight spur, 10 to 13mm long, flower bilateral, snapdragon form, pale yellow with dark orange centre
  • 
<!--Next: FOR BLIND INFLORESENCE ICON. -->
Spike-like 
inflorescence diagram
    • Spike-like 
inflorescence diagram
    • INFLORESCENCE TYPE
    • Spike-like
    • --flowers concentrated at the ends of each stem in an elongated fashion, either erect or drooping.
    • This plant:
    • Tall, upright, one-sided spikes bearing 20-50 flowers
  • 20 to 70 cm
  • Jun Jul Aug Sep Oct
  • Fields, wayside

    Dry scrub

    • For more information, go to the “Habitats & such” tab in the menu bar.
  • diagram of photo location in the Lobau
    • diagram of photo location in the Lobau
Common Toadflax
Linaria vulgaris
Other common name(s): Yellow Toadflax, Butter and Eggs, Wild Snapdragon
German name(s): Echt-Leinkraut, Gewöhnliches Leinkraut, Frauenflachs, Kleines Löwenmaul
ITIS Serial #: 33216
Date:
23 June, 2012

The stems are upright, usually with a one-sided spike of large, pale yellow, spurred flowers, 20 – 50 in number. The spur, 1-1.3cm long, contains nectar and is almost as long as the petals. The underlip has a deep yellow to orange, swollen centre and it closes the entrance to the flower parts so tightly that pollination requires strong insects such as bees and bumblebees (Bombus species) to put sufficient weight on it to activate the spring-like opening. Some short-tongued bees may make a short cut to the nectar by biting a hole in the spur.

The plant is a nectar food plant for a large number of insects.

Bluish-green leaves up to 6cm long are narrow with a distinct mid-rib, the margins slightly rolled under and arranged at close intervals around the stem beneath the flower spike.

Roots may be up to 1 m deep enabling the plant to withstand dry conditions and survive the winter underground. Side shoots develop into daughter plants producing dense groups of genetically identical plants.

The fruit is a capsule with pores. The tiny disc-shaped seeds, up to 3,200 per plant, are shaken out when ripe and distributed further by the wind.


Austrian Speedwell
 in Lobau

© 2014 Margaret Eros

LEAF TYPE
PETAL TYPE
INFLORES- CENCE TYPE
HEIGHT OF PLANT
FLOWERING MONTH
HABITAT
PHOTO LOCATION
  • diagram of 
<!--Next: FOR BLIND LEAF SHAPE ICON.-->
simple 
<!--Next brown section: FOR BLIND LEAF VEIN ICON.-->
 leaf shape and 
pinnate 
vein structure
    • diagram of 
<!--Next: FOR BLIND POPUP LEAF SHAPE.-->
simple 
leaf shape 
<!--Next: FOR BLIND POPUP LEAF VEIN ICON.-->
and 
pinnate 
vein structure
    • LEAF TYPE
    • Shape:
    • --simple (undivided/entire).
    • Veins:
    • --pinnate (main central vein with side branches).
    • This plant:
    • Elongated shape, leaf margin bluntly toothed, leaf stalks (petioles) short or absent
  •  4 separate petals 
flower type diagram
    •  4 separate petals 
flower type diagram
    • FLOWER TYPE
    • Symmetry:
    • Petals:
    • --4.
    • This plant:
    • Light blue, symmetry radial at first view but slightly bilateral when examined closely (see !D!)
  • 
<!--Next: FOR BLIND INFLORESENCE ICON. -->
Spike-like 
inflorescence diagram
    • Spike-like 
inflorescence diagram
    • INFLORESCENCE TYPE
    • Spike-like
    • --flowers concentrated at the ends of each stem in an elongated fashion, either erect or drooping.
    • This plant:
    • Dense spikes of small flowers carried at tips of upwardly-turning horizontal or upright stems.
  • 10 to 40 cm
  • Apr May Jun
  • Dry grassland

    • For more information, go to the “Habitats & such” tab in the menu bar.
  • diagram of photo location in the Lobau
    • diagram of photo location in the Lobau
Austrian Speedwell
Veronica austriaca
Other common name(s): Saw-leaved Speedwell
German name(s): Österreichische Ehrenpreis
ITIS Serial #: 505667
Date:
16 April, 2014
A protected flower.

This is a perennial plant but without stolons (runners) so doesn’t spread laterally. Grows in dense clumps, the stems are weak and radiate in a spreading fashion from the base, sometimes only the flowering tips bend upwards in an upright fashion. This gives the impression of shorter stems than they really are. Nevertheless, they rarely reach the maximum of 40cm. Stem lengths do not normally exceed 25cm in the Lobau.

Blooming usually begins in May but recent warmer Spring weather has pushed forward the flowering time to include April.

Petals are light blue with darker lines; the centre of the flower is white. There are 2 stamens (male organs with dark blue anthers) and a single long style (female organ carrying a stigma at the tip that catches pollen grains).

  • !D!
    • single flower of Austrian speedwell showing slight bilateral symmetry
    • single flower showing lined markings on petals, slight bilateral symmetry, long single style and two stamens
  • !L!
    • elongated, stalkless (sessile) leaves  of Austrian Speedwell
    • elongated, stalkless (sessile) leaves with saw-like toothed margins
  • !W!
    • whole clump of Austrian Speedwell  flowers showing stems radiating from central base
    • Stems radiate from base, outer ones lying flat on ground, turning upwards at the tip

Saw-leaved Speedwell
 in Lobau

© 2014 Margaret Eros

LEAF TYPE
PETAL TYPE
INFLORES- CENCE TYPE
HEIGHT OF PLANT
FLOWERING MONTH
HABITAT
PHOTO LOCATION
  • diagram of 
<!--Next: FOR BLIND LEAF SHAPE ICON.-->
simple 
<!--Next brown section: FOR BLIND LEAF VEIN ICON.-->
 leaf shape and 
pinnate 
vein structure
    • diagram of 
<!--Next: FOR BLIND POPUP LEAF SHAPE.-->
simple 
leaf shape 
<!--Next: FOR BLIND POPUP LEAF VEIN ICON.-->
and 
pinnate 
vein structure
    • LEAF TYPE
    • Shape:
    • --simple (undivided/entire).
    • Veins:
    • --pinnate (main central vein with side branches).
    • This plant:
    • Oval shape, bluntly pointed, toothed margins, arranged in opposite pairs
  • 4 separate petals 
flower type diagram
    • 
4 separate petals 
flower type diagram
    • FLOWER TYPE
    • Symmetry:
    • Petals:
    • --4.
    • This plant:
    • symmetry slightly bilateral, upper petal larger, lower petal narrower. (see !D! button below)
  • 
<!--Next: FOR BLIND INFLORESENCE ICON. -->
Spike-like 
inflorescence diagram
    • 
Spike-like 
inflorescence diagram
    • INFLORESCENCE TYPE
    • Spike-like
    • --flowers concentrated at the ends of each stem in an elongated fashion, either erect or drooping.
    • This plant:
    • young spikes droop, becoming erect as flowers successively open
  • 15 to 80 cm
  • May Jun Jul
  • Woodland edge

    Grassland

    • For more information, go to the 'Habitats & such' tab in the menu bar.
  • diagram of photo location in the Lobau
    • diagram of photo location in the Lobau
Saw-leaved Speedwell
Veronica teucrium; syn Veronica austriaca ssp teucrium
Other common name(s):
German name(s): Grosser Ehrenpreis
ITIS Serial #: 523226
Date:
01 June, 2014

This is a Central and south European species that thrives in sunny locations on the edges of woods in grassy places.

It is a perennial plant that overwinters in the form of underground stems or rhizomes.

The form of the flowers is similar to those of Veronica austriaca as described above but the plant differs in various ways. It grows in grassy woodland edges as opposed to dry grassland areas and is taller with more erect stems. (The Austrian Speedwell is lower growing with prostrate stems). The inflorescence spike is longer and finer, drooping in the early stages before the flowers are fully open. It flowers later than Austrian Speedwell, blooms first appearing in late May, early June. The leaves are broader and thinner textured with a regular dentate margin.

(These identifications require further corroboration and validation. )

  • !D!
    • single flower showing slight bilateral symmetry
    • Single flower showing slight bilateral symmetry
  • !W!
    • long drooping spikes at beginning of flowering
    • Long drooping inflorescence spikes at beginning of flowering
  • !L!
    • Leaves arranged in opposite pairs; dentate margins
    • Leaves arranged in opposite pairs; dentate margins
Persian Speedwell
 in Lobau

© 2011 Margaret Eros

LEAF TYPE
PETAL TYPE
INFLORES- CENCE TYPE
HEIGHT OF PLANT
FLOWERING MONTH
HABITAT
PHOTO LOCATION
  • diagram of 
<!--Next: FOR BLIND LEAF SHAPE ICON.-->
simple 
<!--Next brown section: FOR BLIND LEAF VEIN ICON.-->
 leaf shape and 
pinnate 
vein structure
    • diagram of 
<!--Next: FOR BLIND POPUP LEAF SHAPE.-->
simple 
leaf shape 
<!--Next: FOR BLIND POPUP LEAF VEIN ICON.-->
and 
pinnate 
vein structure
    • LEAF TYPE
    • Shape:
    • --simple (undivided/entire).
    • Veins:
    • --pinnate (main central vein with side branches).
    • This plant:
    • short-stalked leaves broadly ovate with course serrated margins, 1-2cm long, heart-shaped at base
  • Radial symmetry and 4 separate petals 
flower type diagram
    • Radial symmetry and 4 separate petals 
flower type diagram
    • FLOWER TYPE
    • Symmetry:
    • --radial (star-shaped as seen from above).
    • Petals:
    • --4.
    • This plant:
    • symmetry slightly bilateral on closer viewing – one petal different in shape, narrower and more angular, 2 stamens, one long style
  • 
<!--Next: FOR BLIND INFLORESENCE ICON. -->
single 
inflorescence diagram
    • Single 
 inflorescence diagram
    • INFLORESCENCE TYPE
    • Single
    • --each flower stem carries a single solitary flower.
    • This plant:
    • simple branching from axils along prostrate leafy stem, single flower per branch.
  • 10 to 30 cm
  • Mar Apr May Jun Jul Aug Sep Oct Nov
  • Fields

    Wayside

    • For more information, go to the “Habitats & such” tab in the menu bar.
  • diagram of photo location in the Lobau
    • diagram of photo location in the Lobau
Persian Speedwell
Veronica persica
Other common name(s): Bird’s-eye Speedwell
German name(s): Persischen Ehrenpreis
ITIS Serial #: 33405
Date:
11 May, 2011

This is an annual, sometimes overwintering biannual plant, reproducing from seed. It has a weak stem and prostrate growth forming a dense ground cover. Although the stem can reach up to 30cm, it never roots along its length.

The flowers are roughly a centimetre wide and are sky-blue with dark stripes and white centres. Although not evident at first glance, they are slightly zygomorphic (bilaterally symetrical with one vertical plane of symmetry), growing singly on long, slender, hairy stalks in the leaf axils.

  • !?!
    • 
Swallowtail butterfly with title ‘did you know’
    • Thought to have originated in the Caucasus, Persian Speedwell was first brought to Europe to expand flower collections in botanical gardens. At the beginning of the 19th century, it ‘escaped’ and has since spread to become a common wild flower throughout Central Europe.
  • !W!
    • 
tip of growing stem of Persian speedwell showing flowers developing in axils of each leaf
    • tip of growing stem showing flowers developing in axils of each leaf